[1] Univ Roma Tre, Dipartimento Sci, Largo San Leonardo Murialdo 1, I-00146 Rome, Italy
[2] Univ Firenze, Dipartimento Sci Terra, Via G La Pira 4, I-50121 Florence, Italy
[3] Univ Roma La Sapienza, Dipartimento Sci Terra, Piazzale Aldo Moro 5, I-00185 Rome, Italy
[4] Univ New England, Form Evolut & Anat Res Lab, Zool, Sch Environm & Rural Sci, Armidale, NSW 2351, Australia