[1] Univ Western Australia, Sch Anat & Human Biol, Nedlands, WA 6009, Australia
[2] Univ Bern, Fac Med, Dept Clin Res, CH-3012 Bern, Switzerland
[3] Univ Western Australia, Nedlands, WA 6009, Australia
[4] King Edward Mem Hosp, Nedlands, WA 6009, Australia