[1] Aarhus Univ, Arct Res Ctr, Dept Biosci, DK-8000 Aarhus C, Denmark
[2] Norwegian Inst Nat Res, Trondheim, Norway
[3] Smithsonian Environm Res Ctr, Edgewater, MD USA
[4] Norwegian Inst Air Res, Stjordal, Norway
[5] Lund Univ, Dept Phys Geog & Ecosyst Sci, S-22100 Lund, Sweden
[6] Harvard Univ, Dept Organism & Evolutionary Biol, Cambridge, MA 02138 USA
[7] Norwegian Inst Agr & Environm Res, Bioforsk, Stjordal, Norway
[8] Waterford Inst Technol, Dept Sci, Waterford, Ireland
[9] Univ Coll Cork, Environm Res Inst, Cork, Ireland