[1] Inst Invest Marinas & Costeras INVEMAR, Soc Portuaria, Programa Biodiversidad & Ecosistemas Marinos, Santa Marta, Colombia
[2] Univ Cadiz, Fac Ciencias Mar & Ambientales, Dept Geol, Puerto Real 11510, Spain
[3] Univ Catolica Norte, Fac Ciencias Mar, Dept Biol Marina, Larrondo 1281, Coquimbo, Chile
[4] Univ Concepcion, Ctr Invest Oceanog Pacifico S Oriental COPAS, Concepcion, Chile