[1] NOAA, Natl Marine Fisheries Serv, NW Fisheries Sci Ctr, Seattle, WA 98112 USA
[2] NOAA, Protected Resources Off, Natl Marine Fisheries Serv, Silver Spring, MD USA
[3] US Fish & Wildlife Serv, Endangered Species Program, Arlington, VA USA
[4] Dalhousie Univ, Dept Biol, Halifax, NS, Canada
[5] Univ Oslo, Ctr Ecol & Evolutionary Synth, Dept Biosci, N-0316 Oslo, Norway