[1] NOAA, Pacific Isl Fisheries Sci Ctr, Ecosyst Sci Div, Honolulu, HI 96818 USA
[2] Univ Hawaii, Joint Inst Marine & Atmospher Res, Honolulu, HI 96822 USA
[3] NOAA, Pacific Marine Environm Lab, 7600 Sand Point Way Ne, Seattle, WA 98115 USA
[4] Woods Hole Oceanog Inst, Woods Hole, MA 02543 USA
[5] Boston Univ, Boston, MA 02215 USA
[6] Wildlife Conservat Soc, Suva, Fiji
[7] US Fish & Wildlife Serv, Marine Natl Monuments Pacific, Honolulu, HI USA