[1] Univ Louisville, Dept Anat Sci & Neurobiol, Louisville, KY 40292 USA
[2] Museo Argentino Ciencias Nat Bernardino Rivadavia, Dept Vertebrate Paleontol, CONICET, Buenos Aires, DF, Argentina
[3] Univ Nacl La Pampa, CONICET, RA-6300 La Pampa, Argentina
[4] Amer Museum Nat Hist, Div Paleobot, New York, NY 10024 USA
[5] Columbia Univ, Lamont Doherty Earth Observ, Dept Earth & Environm Sci, New York, NY 10964 USA