[1] Univ Utrecht, Dept Phys Geog, NL-3508 TC Utrecht, Netherlands
[2] Univ Utrecht, Fac Vet Med, NL-3584 CL Utrecht, Netherlands
[3] Stavropol Plague Control Res Inst, Stavropol 355035, Russia
[4] M Aikimbayevs Kazakh Sci Ctr Quarantine & Zoonot, Antiplague Inst, Alma Ata 050074, Kazakhstan
[5] RMIT Univ, Sch Math & Geospatial Sci, Melbourne, Vic 3000, Australia
[6] Univ Liverpool, Inst Integrat Biol, Liverpool L69 3BX, Merseyside, England
[7] Univ Antwerp, Dept Biol, B-2020 Antwerp, Belgium