The Evolution of Tribalism: A Social-Ecological Model of Cooperation and Inter-group Conflict under Pastoralism

This study investigates a possible nexus between inter-group competition and intra-group cooperation, which may be called "tribalism." Building upon previous studies demonstrating a relationship between the environment and social relations, the present research incorporates a social-ecological model as a mediating factor connecting both individuals and communities to the environment. Cyclical and non-cyclical fluctuation in a simple, two-resource ecology drive agents to adopt either "go-it-alone" or group-based survival strategies via evolutionary selection. Novelly, this simulation employs a multilevel selection model allowing group-level dynamics to exert downward selective pressures on individuals' propensity to cooperate within groups. Results suggest that cooperation and inter-group conflict are co-evolved in a triadic relationship with the environment. Resource scarcity increases inter-group competition, especially when resources are clustered as opposed to widely distributed. Moreover, the tactical advantage of cooperation in the securing of clustered resources enhanced selective pressure on cooperation, even if that implies increased individual mortality for the most altruistic warriors. Troubling, these results suggest that extreme weather, possibly as a result of climate change, could exacerbate conflict in sensitive, weather-dependent social-ecologies - especially places like the Horn of Africa where ecologically sensitive economic modalities overlap with high-levels of diversity and the wide- availability of small arms. As well, global development and foreign aid strategists should consider how plans may increase the value of particular locations where community resources are built or aid is distributed, potentially instigating tribal conflict. In sum, these factors, interacting with pre-existing social dynamics dynamics, may heighten inter-ethnic or tribal conflict in pluralistic but otherwise peaceful communities.