[1] Univ Cote Azur, CNRS, ECOSEAS, Lyon, France
[2] Lab Int Associe Univ Cote Azur, Ctr Scienti fi que Monaco, LIA ROPSE, Nice, Monaco
[3] Sorbonne Univ, CNRS, UMR 7093, LOV, Villefranche sur Mer, France
[4] Stn Zool Anton Dohrn Natl Inst Marine Biol Ecol &, Genoa Marine Ctr, Dept Integrat Marine Ecol EMI, Genoa, Italy
[5] Ctr Sci Monaco, Monaco, Monaco