[1] Field Museum, Bot Dept, Chicago, IL 60605 USA
[2] Univ Nacl Agr La Molina, Herbario Forestal, Lima, Peru
[3] Eotvos Lorand Univ, Dept Plant Systemat Ecol & Theoret Biol, Budapest, Hungary
[4] Univ Hohenheim, Dept Mol Bot, Stuttgart, Germany
[5] Eszterhazy Karoly Catholic Univ, Inst Biol, Bot Dept, Eger, Hungary