[1] Middlesex Univ, Dept Nat Sci, London, England
[2] Univ Fed Pernambuco, Ctr Biociencias, Dept Zool, Lab Ecol Comportamento & Conservacao LECC, Recife, Brazil
[3] Inst Nacl de Pesquisas da Amazonia, Grp Pesquisa Mamiferos Amazonia, Manaus, AM, Brazil
[4] Rhodes Coll, Dept Biol, Memphis, TN USA
[5] Univ Fed Pernambuco, Ctr Biociencias, Dept Zool, Programa Posgrad Biol Anim, Recife, Brazil