[1] Shenyang Normal Univ, Coll Paleontol, Shenyang, Peoples R China
[2] Minist Nat Resources, Key Lab Evolut Life NE Asia, Shenyang, Peoples R China
[3] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, Nanjing, Peoples R China
[4] Univ Gdansk, Fac Biol, Dept Invertebrate Zool & Parasitol, Lab Evolutionary Entomol, Gdansk, Poland
[5] Univ Gdansk, Fac Biol, Dept Invertebrate Zool & Parasitol, Museum Amber Inclus, Gdansk, Poland
[6] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, Nanjing 210008, Peoples R China