[1] South African Natl Biodivers Inst, Compton Herbarium, Private Bag X7, ZA-7735 Cape Town, South Africa
[2] Univ KwaZulu Natal, Res Ctr Plant Growth & Dev, Sch Biol & Conservat Sci, Private Bag X1, ZA-3209 Scottsville, South Africa
[3] POB 22652, ZA-7975 Scarborough, South Africa
[4] Stellenbosch Univ, Dept Bot & Zool, Private Bag X1, ZA-7602 Matieland, South Africa
[5] Univ Cape Town, Dept Biol Sci, Bolus Herbarium, Private Bag X3, ZA-7701 Rondebosch, South Africa