[1] Univ Cologne, Inst Genet, Cologne, Germany
[2] Cluster Excellence Cellular Stress Responses Agin, Cologne, Germany
[3] Max Planck Inst Biol Ageing, Cologne, Germany
[4] Univ Cologne, Dept Neurol, Cologne, Germany
[5] Univ Cologne, Ctr Mol Med CMMC, Cologne, Germany