[1] Univ Minnesota, Dept Plant & Microbial Biol, St Paul, MN 55455 USA
[2] Manchester Metropolitan Univ, Dept Nat Sci, Manchester, England
[3] Natl Ctr Biol Sci, Wildlife Biol & Conservat Program, Bangalore, India
[4] Nanyang Technol Univ, Asian Sch Environm, Singapore, Singapore