[1] Department of Geography and Environmental Science, University of Reading, Whiteknights, Reading,RG6 6AB, United Kingdom
[2] UK Centre for Ecology and Hydrology, MacLean Building, Benson Lane, Crowmarsh Gifford, Wallingford,OX10 8BB, United Kingdom
[3] School of Biological Sciences, University of Reading, Whiteknights, Reading,RG6 6AB, United Kingdom
[4] Centre for Environment, Fisheries & Aquaculture Science, Lowestoft Laboratory, Pakefield Road, Lowestoft,NR330HT, United Kingdom
[5] School of Environmental Sciences, University of East Anglia, Norwich Research Park, Norwich,Norfolk,NR4 7TJ, United Kingdom