[1] Niigata Univ, Brain Res Inst, Niigata, Japan
[2] Tokyo Inst Technol, Sch Life Sci & Technol, Yokohama, Japan
[3] Yokohama Natl Univ, Res Initiat & Promot Org, Yokohama, Japan
[4] Tokyo Metropolitan Univ, Grad Sch Sci, Dept Biol Sci, Hachioji, Japan
[5] Natl Inst Genet, Dept Gene Funct & Phenom, Mishima, Japan
[6] Niigata Univ, Grad Sch Med & Dent Sci, Div Ophthalmol & Visual Sci, Niigata, Japan