[1] Beijing Forestry Univ, State Key Lab Efficient Prod Forest Resources, Beijing 100083, Peoples R China
[2] Beijing Forestry Univ, Res Ctr Forest Management Engn State Forestry & Gr, Beijing 100083, Peoples R China
[3] Jilin Prov Acad Forestry Sci, Changchun 130033, Peoples R China
[4] Georg August Univ Gottingen, Fac Forestry & Forest Ecol, Gottingen, Germany
[5] Univ Stellenbosch, Dept Forest & Wood Sci, Stellenbosch, South Africa