[1] Block 207A,Woodleigh Link, Singapore 361207, Singapore
[2] PSL, Inst Systemat Evolut Biodivers ISYEB, Museum Natl Hist Nat, CNRS,EPHE,SU,UA, 57 Rue Cuvier,CP 50, F-75231 Paris 05, France
[3] Univ Malaya, Inst Biol Sci, Fac Sci, Kuala Lumpur 50603, Malaysia
[4] Forest Res Ctr Sepilok, Sabah Forestry Dept, POB 1407, Sandakan 90715, Sabah, Malaysia