[1] Univ British Columbia, Dept Wood Sci, Vancouver, BC V6T 1Z4, Canada
[2] York Timbers, Res Div, Sabie, Mpumalanga, South Africa
[3] York Timbers, Planning Div, Sabie, Mpumalanga, South Africa
[4] Stellenbosch Univ, Dept Forestry, Stellenbosch, Western Cape, South Africa
[5] York Timbers, Tree Breeding Div, Sabie, Mpumalanga, South Africa
[6] Univ Witwatersrand, Fac Sci, Sch Anim Plant & Environm Sci, Johannesburg, Gauteng, South Africa
[7] Inst Commercial Forestry Res, Pietermaritzburg, Kzn, South Africa
[8] Univ Pretoria, Dept Biochem Genet & Microbiol, Pretoria, Gauteng, South Africa
[9] CharlieC Consulting, Cheltenham GL54 4NU, England
[10] Univ British Columbia, Dept Bot, Vancouver, BC V6T 1Z4, Canada