[1] Univ Lyon, Inst Syst Complexes 9011, 46 Allee Italie, F-69364 Lyon 07, France
[2] Univ Lyon, Ecole Normale Super Lyon, Lab Reprod & Dev Plantes, INRAE,UCB Lyon 1,INRAE,CNRS, 46 Allee Italie, F-69364 Lyon 07, France
[3] European Environm Agcy, Sci Comm, Frankrigshusene 9,1 tv, DK-2300 Copenhagen S, Denmark