[1] Univ Granada, Fac Sci, Dept Ecol, Ave Fuente Nueva S-N, Granada 18071, Spain
[2] Univ Granada, Andalusian Inst Earth Syst Res IISTA CEAMA, Lab Ecol, Ave Mediterraneo S-N, Granada 18006, Spain
[3] Environm & Water Agcy, Dept Sustainabil Environm & Blue Econ Reg Govt And, C Minerva 7 Edificio Zeus 3, Granada 18014, Spain
[4] Spanish Natl Res Council IEO CSIC, Spanish Inst Oceanog, St ander Oceanog Ctr COST IEO, Ave Severiano Ballesteros 16, Santander 39004, Spain
[5] Univ Granada, Fac Sci, Dept Zool, Ave Fuente Nueva S-N, Granada 18071, Spain