[1] Univ Sains Malaysia, Sch Biol Sci, Usm Pulau Pinang 11800, Malaysia
[2] Univ Sains Malaysia, Ctr Marine & Coastal Studies CEMACS, Usm Pulau Pinang 11800, Malaysia
[3] Int Islamic Univ Malaysia, Dept Marine Sci, Kulliyyah Sci, Kuantan 25200, Pahang, Malaysia
[4] Int Islamic Univ Malaysia, Inst Oceanog & Maritime Studies INOCEM, Kulliyyah Sci, Kuantan 25200, Pahang, Malaysia